Tohoku J. Exp. Med., 221: 287-298, 2010. In the version of this article published in the August issue, 2010, the affiliation number of the tenth author (Shigeru Tsuchiya) was mistyped (page 287). The correct affiliation is shown above. In addition, the lettering of ‘ZT’ is a misprint for ‘CT’ in panels A, B and C of Figures 6 and 7 (pages 294 and 295).