Paracytic (Syndetocheilic) Stomata in Carboniferous Seed Ferns

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Abstract
Stomata of the paracytic type have been discovered on the lower pinnule surface of the Carboniferous seed fern foliage species Alethopteris sullivanti . The stomatal complex is not visible when the cuticle is in place. This is apparently the first report of paracytic stomata in Carboniferous plants.