Spherical squirmers: models for swimming micro-organisms

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Abstract
In 1952, Lighthill introduced the simplest possible model of a swimming micro-organism of finite size, intended as a model of a single-celled protozoan covered in beating cilia. The model consisted of a sphere, on the surface of which material points undergo small-amplitude oscillations. In 1971, Lighthill’s student, John Blake, completed the calculations and applied the theory to various species of ciliate. Subsequently we, and many others, have used the even simpler (though less realistic) model of a steady squirmer, a sphere whose surface moves tangentially with time-independent velocity. In this article we survey:

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