Kevin Paul Kenow

In cooperation with the Minnesota Department of Natural Resources, the U.S. Geological Survey monitored 98 Gavia immer (common loon) focal territories and 43 nonfocal territories in north-central Minnesota in 2025. Focal territories are those sampling units from which study inferences can be made. Nonfocal territories were adjacent to focal territories and were observed to monitor common loon dynamics across an entire waterbody. In collaboration with lake associations and private citizens, we deployed 44 artificial nesting platforms to 44 focal territories, called treatment territories. Territorial surveys to monitor occupancy, nesting, and chick survival were completed between April 27 and August 14, 2025. We attempted to visit each focal and nonfocal territory once per week. At least 1 nest attempt was observed in 40 of 54 control territories and 37 of 44 treatment territories. In treatment territories, 14 nests were on an artificial nesting platform, and the remaining nests were on natural substrates. Chicks or other evidence of hatching were observed in 20 of 54 control territories (37 percent) and 28 of 44 treatment territories (64 percent). Twelve of those successful nests in treatment territories were on artificial nesting platforms.

In cooperation with the Minnesota Department of Natural Resources, the U.S. Geological Survey monitored 98 common loon (Gavia immer) focal territories and an additional 37 nonfocal territories in 2024 across 53 study lakes in Minnesota. Focal territories were those territories from which study inferences will be made, whereas nonfocal territories were observed to monitor common loon dynamics in territories adjacent to focal territories. In collaboration with lake associations and private citizens, we deployed 44 artificial nesting platforms within 44 treatment territories, and the remaining 54 focal territories were controls. We completed territorial surveys from April 29 to August 9, 2024, to evaluate occupancy, nest success, and chick survival. We attempted to visit each territory once a week. At least one nest attempt was observed in 41 of 54 control territories. Precisely one nest attempt was observed in 30 control territories, two nest attempts (first attempts failed) were observed in 9 control territories, and three nest attempts (first and second attempts failed) were observed in 2 control territories. At least one nest attempt was observed in 33 of 44 treatment territories. Precisely one nest attempt was observed in 25 treatment territories, two nest attempts (first attempts failed) were observed in 7 treatment territories, and three nest attempts (first and second attempts failed) were observed in 1 treatment territory. In treatment territories, 8 nests were on an artificial nesting platform; the remaining nest locations were natural. Chicks or other evidence of hatching were observed in 26 of 54 (48.1 percent) control territories and 21 of 44 (47.7 percent) treatment territories, with 7 of those successful treatment nests on an artificial nesting platform. This report includes no formal analysis, but we plan to analyze data after collection of all field data in subsequent years.

The Deepwater Horizon mobile drilling platform exploded on April 20, 2010. The resulting massive oil spill injured natural resources in the Gulf of Mexico, including wintering common loons (Gavia immer). We report on activities completed under the “Restoration of Common Loons in Minnesota” project in calendar year 2023, which was funded by the Open Ocean Trustee Implementation Group. In 2022, a subset of monitored breeding territories was identified as focal territories, which are sampling units for the study. The U.S. Geological Survey, in cooperation with the Minnesota Department of Natural Resources, monitored 98 common loon focal territories and an additional 43 nonfocal territories in 2023 across 56 study lakes in Minnesota. We collaborated with lake associations and private citizens to deploy 42 artificial nesting platforms within 44 focal treatment territories. The remaining 54 focal territories were controls. Territorial surveys were completed from May 8 to August 11, 2023, to evaluate occupancy, nest success, and chick survival. At least one nest attempt was observed in 31 of 44 treatment territories and a second nest attempt was observed after a failed initial attempt in 6 treatment territories. However, only one nest was on an artificial nesting platform in a treatment territory; the remaining nest locations were natural. At least one nest attempt was observed in 37 of 54 control territories, and a second nest attempt was observed after a failed initial attempt in 5 control territories. Chicks or other evidence of hatching were observed in 17 of 54 control territories and 17 of 44 treatment territories, with 1 of those successful treatment nests occurring on an artificial nesting platform. This report includes no formal analysis, but we plan to analyze data after collection of all field data in subsequent years.

Common loon ( Gavia immer (Brünnich, 1764)) foraging patterns and the relative importance of cisco ( Coregonus artedi Lesueur, 1818) in the diets of loons were evaluated for the Whitefish Chain of Lakes, a set of coldwater cisco refuge lakes in Minnesota, USA. Environmental DNA metabarcoding of loon fecal samples detected 15 fish species. Yellow perch ( Perca flavescens (Mitchill, 1814)), mimic shiner ( Notropis volucellus (Cope, 1865)), and cisco were the most prominent prey detected across the study lakes. We observed a shift in fish species consumed, with increases in detections of mimic shiner and cisco DNA among loon fecal samples collected in August and September. In some locations, suitable oxythermal habitat became restricted throughout the summer, forcing cisco into surface waters, which potentially increased their vulnerability to loon predation. Conversely, large foraging aggregations of loons were observed during late summer through fall at locations with ample oxythermal habitat and abundant cisco populations. We hypothesize that cisco were sought by loons as a high-calorie prey resource prior to migration. Conservation efforts directed at preserving water quality in important cisco refuge lakes are likely to benefit common loons through enhancement of both the forage base, for resident and migrating birds, and breeding habitat suitability.

The explosion of the Deepwater Horizon mobile drilling platform on April 20, 2010, caused a massive oil spill and injury to natural resources in the Gulf of Mexico. Gavia immer (common loon) were negatively affected from the spill. The Open Ocean Trustee Implementation Group funded the project “Restoration of Common Loons in Minnesota” to restore common loons lost to the spill. Here, we report on activities conducted for this project in an eight-county region in Minnesota in calendar year 2022. We identified a subset of territories that were monitored in 2021 as focal territories (n=99) from which multiyear study inferences can ultimately be made. We monitored nonfocal territories on all study lakes as well. We conducted surveys from May 9 to August 12, 2022. At least 1 nest attempt was observed in 68 of 99 focal territories, and a second nest attempt after a failed initial attempt was observed in 19 focal territories. Chicks or other evidence of hatching was observed in 33 of 99 territories. Data collected in 2021 and 2022 for this project are presented in a U.S. Geological Survey data release (https://doi.org/10.5066/P9LA536E). We present no formal data analysis in this report.

Common Loons (Gavia immer) winter primarily in marine coastal areas and utilize a forage base that is poorly defined, especially for offshore areas. Information on dive activity is needed for describing foraging strategies and for inferring prey distribution. Archival geolocator tags were used to determine the wintering locations and dive characteristics of adult Common Loons captured and marked on breeding lakes in Minnesota, Wisconsin, and the Upper Peninsula of Michigan. Among loons that completed fall migration, most wintered in the Gulf of Mexico, with smaller proportions wintering off the southern Atlantic Coast or impoundments in the southeastern United States. Adult Common Loons tended to occupy offshore areas of the Gulf of Mexico and the Atlantic Ocean and, on average, spent about 60% of daylight hours foraging. Dive depths were as deep as 50 m (Gulf of Mexico) and dive characteristics indicated that loons were primarily foraging on benthic prey. Total dive duration, time at maximum depth, and postdive surface intervals increased with dive depths among wintering Common Loons. Our results are expected to contribute to the understanding of the wintering ecology of Common Loons and be useful in informing regional and national conservation planning efforts. RESUMEN. Los colimbos comunes (Gavia immer) invernan principalmente en zonas costeras marinas y utilizan una base para su alimentación que está mal definida, especialmente en las zonas de altamar. La información sobre la actividad de buceo es necesaria para describir las estrategias de alimentación y para inferir la distribución de las presas. Se utilizaron marcadores geolocalizadores de archivo para determinar los lugares de invernada y las características de inmersión de los colimbos comunes adultos capturados y marcados en lagos de cría de Minnesota, Wisconsin y la Península Superior de Michigan. Entre los colimbos que completaron la migración otoñal, la mayoría invernó en el Golfo de México, y en menor proporción en la costa atlántica meridional o en embalses del sureste de Estados Unidos. Los adultos de colimbo común tienden a ocupar zonas costeras del Golfo de México y el Océano Atlántico y, en promedio, pasan alrededor del 60% de las horas de luz buscando alimento. Las profundidades de inmersión alcanzaron los 50 m (Golfo de México) y las características de la inmersión indicaron que los colimbos se alimentaban principalmente de presas bentónicas. La duración total de la inmersión, el tiempo a máxima profundidad y los intervalos en superficie tras la inmersión aumentaron con la profundidad de la inmersión entre los colimbos invernantes. Se espera que nuestros resultados contribuyan a la comprensión de la ecología de invernada de los colimbos comunes y sean útiles para informar los esfuerzos regionales y nacionales de planificación de la conservación.

The Deepwater Horizon oil spill caused extensive injury to natural resources in the Gulf of Mexico, and Gavia immer (common loon) were negatively affected from the spill. The Open Ocean Trustee Implementation Group funded the project Restoration of Common Loons in Minnesota to restore common loons lost to the spill. In 2020–21, priority lakes in an eight-county region in north-central Minnesota were identified to focus project activities. In 2021, surveys on these lakes were started to monitor common loon territory occupancy, nest success, and chick survival. We surveyed 62 lakes and identified 110 common loon territories that will be included in the project. At least 1 nest attempt was observed in 78 of 110 territories, and a second nest attempt was observed in 23 territories. A third nest attempt was observed in one territory. Successful nesting was observed in 32 of 110 territories. We present no formal data analysis and plan to analyze the data after the collection of all field data in subsequent years.

Identification of geographic linkages among breeding, migratory and wintering common loon Gavia immer populations is needed to inform regional and national conservation planning efforts and compensation of loons lost during marine oil spill events. Satellite telemetry and archival geolocator tags were used to determine the migration patterns and wintering locations of breeding adult and young of the year juvenile common loons captured and marked on lakes in Minnesota, Wisconsin and Michigan. Adult loons typically traveled from breeding lakes, often via larger staging lakes, to the Great Lakes (primarily Lake Michigan) and then on to wintering areas. Most radiomarked juvenile common loons utilized natal lakes or local lakes through mid‐November. Subsequently, the first fall migration of juvenile loons was generally initiated later, and more direct and quicker to wintering areas relative to adults. Among adult (n = 103) and juvenile (n = 23) loons that completed fall migration, most wintered in the Gulf of Mexico (GOM), with smaller proportions wintering off the southern Atlantic Coast or impoundments in the southeastern United States. Spring migration of adults to breeding lakes was less prolonged than fall migration, with adult male loons tending to depart wintering areas earlier than adult females. Juvenile common loons migrated during their first spring from wintering sites in the GOM to summer in the Gulf of St Lawrence/Nova Scotia Coastal region. Juvenile mortality was largely linked to parasitic infection and emaciation; spring appeared to be a survival bottleneck among juvenile loons monitored in our study. Our results identify several areas where common loon conservation efforts could be directed to protect key habitats and minimize stressors during the non‐breeding period.

Long‐distance migration presents complex conservation challenges, and migratory species often experience shortfalls in conservation due to the difficulty of identifying important locations and resources throughout the annual cycle. In order to prioritize habitats for conservation of migratory wildlife, it is necessary to understand how habitat needs change throughout the annual cycle, as well as to identify key habitat sites and features that concentrate large numbers of individuals and species. Among long‐distance migrants, sea ducks have particularly complex migratory patterns, which often include distinct post‐breeding molt sites as well as breeding, staging and wintering locations. Using a large set of individual tracking data (n = 476 individuals) from five species of sea ducks in eastern North America, we evaluated multi‐species habitat suitability and partitioning across the breeding, post‐breeding migration and molt, wintering and pre‐breeding migration seasons. During breeding, species generally occupied distinct habitat areas, with the highest levels of multi‐species overlap occurring in the Barrenlands west of Hudson Bay. Species generally preferred flatter areas closer to lakes with lower maximum temperatures relative to average conditions, but varied in distance to shore, elevation and precipitation. During non‐breeding, species overlapped extensively during winter but diverged during migration. All species preferred shallow‐water, nearshore habitats with high productivity, but varied in their relationships to salinity, temperature and bottom slope. Sea ducks selected most strongly for preferred habitats during post‐breeding migration, with high partitioning among species; however, both selection and partitioning were weaker during pre‐breeding migration. The addition of tidal current velocity, aquatic vegetation presence and bottom substrate improved non‐breeding habitat models where available. Our results highlight the utility of multi‐species, annual‐cycle habitat assessments in identifying key habitat features and periods of vulnerability in order to optimize conservation strategies for migratory wildlife.

Hotspot analysis is a commonly used method in ecology and conservation to identify areas of high biodiversity or conservation concern. However, delineating and mapping hotspots is subjective and various approaches can lead to different conclusions with regard to the classification of particular areas as hotspots, complicating long‐term conservation planning. We present a comparative analysis of recent approaches for identifying waterbird hotspots, with the goal of developing insights about the appropriate use of these methods. We selected four commonly used measures to identify persistent areas of high use: kernel density estimation, Getis‐Ord G_i*, hotspot persistence and hotspots conditional on presence, which represent the range of quantitative hotspot estimation approaches used in waterbird analyses. We applied each of the methods to aerial survey waterbird count data collected in the Great Lakes from 2012–2014. For each approach, we identified areas of high use for seven species/species groups and then compared the results across all methods and to mean effort‐corrected counts. Our results indicate that formal hotspot analysis frameworks do not always lead to the same conclusions. The kernel density and Getis‐Ord G_i* methods yielded the most similar results across all species analysed and were generally correlated with mean effort‐corrected count data. We found that these two models can differ substantially from the hotspot persistence and hotspots conditional on presence estimation approaches, which were not consistently similar to one another. The hotspot persistence approach differed most significantly from the other methods but is the only method to explicitly account for temporal variation. We recommend considering the ecological question and scale of conservation or management activities prior to designing survey methodologies. Deciding the appropriate definition and scale for analysis is critical for interpretation of hotspot analysis results as is inclusion of important covariates. Combining hotspot analysis methods using an integrative approach, either within a single analysis or post hoc, could lead to greater consistency in the identification of waterbird hotspots.